Lycopsis

Lycopsis Temporal range: Mid-Late Miocene (Santacrucian-Chasicoan) ~16.3–9.0 Ma PreꞒ Ꞓ O S D C P T J K Pg N
Fossil of L. longirostrus
Scientific classification
Kingdom:	Animalia
Phylum:	Chordata
Class:	Mammalia
Order:	†Sparassodonta
Superfamily:	†Borhyaenoidea
Genus:	†Lycopsis Cabrera 1927
Species
L. torresi Cabrera 1927 (type species) L. longirostrus Marshall 1976 L. viverensis Forasiepi et al. 2003
Synonyms
Anatherium oxyrhynchus Ameghino 1894

Lycopsis is an extinct genus of South American metatherian that lived during the Miocene in Argentina and Colombia.^[1]

History

Although not named until 1927, Florentino Ameghino described a species now seen as synonymous with Lycopsis torresi, Anatherium oxyrhynchus, in 1895 based on a mandibular ramus with several teeth.^[2][3] The fossil was recovered from Puesto Estancia La Costa in Santa Cruz, Argentina, dating to the Miocene.^[3] The type material of Lycopsis was collected in July 1895 by "C. Berry" from the middle Miocene strata of the Santa Cruz Formation along the Santa Cruz River in the same area.^[2][4] The fossils (MLP 11–113) were fragmentary, constituting only several fragmentary jaw sections from the maxilla and mandible, including several molars.^[4][5] However, these fossils were not named until in 1927, Ángel Cabrera named Lycopsis torresi, the generic name meaning "wolf-like aspect" after the anatomy of the mandible and the specific name after Argentine paleontologist and the director of the Museo de la Plata at the time, Luis Maria Torres.^[4]

Taxonomy

The cladogram after the analysis of Suárez et al., 2015, looks as follows:^[6]

Distribution

Fossils of Lycopsis have been found in:^[3]

• Arroyo Chasicó Formation and Santa Cruz Formations, Argentina
• Honda Group, Colombia

References

• v
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Metatheria

Kingdom: Animalia Phylum: Chordata Class: Mammalia Subclass: Theria
Synapsida see Synapsida Cynodontia see Cynodontia Mammalia see Mammalia Metatheria see below↓

Metatheria
Metatheria †Adinodon †Archaeonothos †Esteslestes †Ghamidtherium †Holoclemensia? †Kasserinotherium †Protalphadon †Sinodelphys? †Deltatheroida Pappotherium? Deltatheridiidae Atokatheridium? Deltatheridium Deltatheroides Gurbanodelta Lotheridium Nanocuris Oklatheridium? Prodeltitheridium Sulestes Tsagandelta Marsupialiformes †Adelodelphys †Aenigmadelphys †Anchistodelphys †Andinodelphys †Arcantiodelphys †Camptomus? †Glasbius †Gurlin Tsav skull †Iugomortiferum †Jaskhadelphys †Mayulestes †Pucadelphys †Sinbadelphys †Sparassodonta see below↓ †Asiatheriidae Asiatherium Kokopellia †Archimetatheria Iqualadelphis Stagodontidae Didelphodon Eobrasilia? Eodelphis Fumodelphodon Hoodootherium Pariadens? Pediomyoidea Aquiladelphis Austropediomys Dakotadens Monodelphopsis Pediomys Protolambda Scalaridelphys Unnuakomys †Peradectidae Armintodelphys Didelphidectes Mimoperadectes Nanodelphys Peradectes Siamoperadectes †Alphadontidae Albertatherium Alphadon Eoalphadon Turgidodon †Herpetotheriidae Amphiperatherium Asiadidelphis Copedelphys Ectocentrocristus Entomacodon Galatiadelphys Golerdelphys Herpetotherium Indodelphis Maastrichtidelphys Morotodon? Nortedelphys Peratherium Rumiodon? Swaindelphys Thylacodon †Anatoliadelphyidae Anatoliadelphys Orhaniyeia †Polydolopimorphia see below↓ Marsupialia Ameridelphia Didelphimorphia Paucituberculata Australidelphia

Sparassodonta
Sparassodonta Allqokirus Argyrolestes Hondadelphys Nemolestes Patene Procladosictis Stylocynus Hathliacynidae Acyon Australogale Borhyaenidium Chasicostylus Cladosictis Notictis Notocynus Notogale Perathereutes Pseudonotictis Sallacyon Sipalocyon Borhyaenoidea Angelocabrerus Chlorocyon Dukecynus Fredszalaya Lycopsis Pharsophorus Plesiofelis Prothylacinus Pseudolycopsis Pseudothylacynus Borhyaenidae Acrocyon Arctodictis Australohyaena Borhyaena Eutemnodus? Proborhyaenidae Arminiheringia Callistoe Paraborhyaena Proborhyaena Thylacosmilidae Anachlysictis Eomakhaira Patagosmilus Thylacosmilus

Polydolopimorphia

Polydolopimorphia Apeirodon? Bobbschaefferia Caroloameghinia Carolocoutoia Guggenheimia Gashternia Hatcheritherium Palangania Procaroloameghinia Protodidelphis Pujatodon? Reigia Robertbutleria Wamradolops Zeusdelphys Bonapartheriiformes Bonapartheriidae Bonapartherium Epidolops Argyrolagidae Anargyrolagus Argyrolagus Hondalagus Microtragulus Proargyrolagus Chulpasiinae Chulpasia Thylacotinga Prepidolopidae Incadolops Perrodelphys Prepidolops Punadolops Seumandia Rosendolopidae Hondonadia Rosendolops Polydolopiformes Polydolopidae Amphidolops Antarctodolops Eudolops Eurydolops Hypodolops Kramadolops Parabderites Polydolops Pseudolops Sillustaniidae Roberthoffstetteria Sillustania